| 1984 | B.A. Washington University, St. Louis, MO, magna cum laude |
| 1984 | M.A. Washington University, St. Louis, MO (Advisor: John P. Atkinson, M.D.) |
| 1990 | Ph.D. Case Western Reserve University (Advisor: Alan M. Tartakoff, Ph.D.) |
| 1991 | M.D. Case Western Reserve University |
| 1991-1993 | Resident in Anatomic Pathology, Brigham and Women’s Hospital |
| 1993 | Fellow in Surgical Pathology (Joseph M. Corson, M.D.), Brigham and Women’s Hospital |
| 1993 | Chief Resident in Anatomic Pathology (Ramzi S. Cotran, M.D.), Brigham and Women’s Hospital |
| 1994-1995 | Fellow in Gastrointestinal and Hepatobiliary Pathology, Brigham and Women’s Hospital |
| 1994-1995 | Research Fellow, Department of Pathology (James L. Madara, M.D.), Brigham and Women’s Hospital |
| 2014 | Sabbatical, Laboratory of Host Defense (Warren Strober, M.D.). National Institute of Allergy and Infectious Disease |
| 1995-1996 | Instructor in Pathology, Brigham and Women’s Hospital and Harvard Medical School |
| 1996-2001 | Assistant/Associate Professor of Pathology, Wayne State University School of Medicine |
| 2001-2016 | Assistant/Associate/Professor of Pathology and Medicine (GI), The University of Chicago |
| 2003-2007 | Associate Residency Director, Department of Pathology, The University of Chicago |
| 2007-2016 | Associate Chair, Department of Pathology, The University of Chicago |
| 2012-2016 | Sara and Harold Lincoln Thompson Professor, The University of Chicago |
| 2016- | Senior Pathologist, Departments of Pathology and Medicine (GI), Brigham and Women’s Hospital |
| 2017- | Professor of Pathology and Medicine, Harvard Medical School |
| 2004-2006 | Associate Editor, Laboratory Investigation |
| 2006-2008 | Senior Associate Editor, Laboratory Investigation |
| 2011-2014 | Associate Editor, Gastroenterology |
| 2013-2014 | Associate Editor, American Journal of Pathology |
| 2014-2019 | Editor-in-Chief (founding), Cellular and Molecular Gastroenterology and Hepatology |